- An axis to which a carpel of a compound pistil may be
attached, as in the case of the geranium; or which is left when a pod
opens. - A columnlike axis in the capsules of mosses. - A term applied to various columnlike parts; as, the
columella, or epipterygoid bone, in the skull of many lizards; the
columella of the ear, the bony or cartilaginous rod connecting the
tympanic membrane with the internal ear. - The upright pillar in the axis of most univalve shells. - The central pillar or axis of the calicles of certain
corals.